Lucille, who goes by the title of conductor, chanteur, and composer for the Royal Orchestra, possesses a unique singing voice that can both purify and destroy Guignols while restoring their memories briefly before they die as humans rather than zombies. Despite his striking good looks, he is often mistaken for a woman due to his somewhat androgynous appearance. This leads to various awkward situations but also provides him with an opportunity to disguise himself. Although Lucille appears to be cheerful and playful most of the time, it's evident that he has experienced a dark past, which is hinted at through flashbacks during gloomy moments. His relationship with Queen Gemsilika is complex and unclear at first, but as the story progresses, it becomes clear that Lucille is a royal offspring with his own mission to fulfill. He struggles with trusting others, likely due to past experiences.